1884 L. M. UnderwoodDescriptive Catal. N. Amer. Hepaticae 43 Elaters [of Aitonia erythrospermæ] quadrispiral.
1997 Jrnl. Paleontol.71 2/2 The megalospheric specimens described here differ from..other species of Polyepidina in possessing a quadrispiral nuclear apparatus.