In the development of distinct antheridial and oogonial cells the allied Ancylistineae show close alliances to Pythium and the Oomycetes.

To study them in their natural position, thin longitudinal sections of the antheridial branch should be made.

Fructification: ovate spores and tufts of antheridial cells attached to the lateral ramuli, which consist of minute, radiating, dichotomous beaded filaments.