the hollow horny barrel of a feather
any of the large stiff feathers of a bird's wing or tail
any of the hollow sharp spines of a porcupine, hedgehog, etc
a bird's feather fashioned into a writing pen
the spasmodic scratching of a quill pen — Hardy
a bobbin, spool, or spindle on which yarn is wound
a hollow shaft often surrounding another shaft and used in various mechanical devices
a roll of dried bark
cinnamon quills
[Middle English quil hollow reed, bobbin, prob from early Low German quiele]