In mosses: (of the operculum or lid of a sporangium) remaining fixed to the columella.
Origin
Late 19th century. From scientific Latin Systylium, genus name and systylius, former specific epithet (1845 or earlier; both from ancient Greek συν- + στῦλος column, with the addition of -ium or -ius, suffix forming adjectives, respectively) + -ous.